PARIS (AFP) - The warring parties in World War I signed no less than 16 peace treaties at the end of the devastating conflict, the most important being the 1919 Treaty of Versailles in which the victorious Allies made Germany accept responsibility.

Here are the main accords.

Brest-Litovsk: March 3, 1918

The first peace treaty of the conflict, it was agreed between Russia - which had sided with the Allies - and Germany and other members of its Central Powers coalition.

Inked in the German-occupied city of Brest-Litovsk, today's Brest in Belarus, it followed a December 1917 armistice sought by an overwhelmed Russia immediately after its October 1917 Marxist revolution.

The terms were harsh: Russia lost to Germany large parts of its western territory in the Baltic states, Finland and Poland, and more than 30 per cent of its population.

Versailles: June 28, 1919

Signed in the mythic Hall of Mirrors at the Palace of Versailles outside Paris exactly five years after the assassination of Archduke Franz Ferdinand that sparked the war, this treaty designated Germany responsible for losses and damages from the war.

Among the terms, Germany was ordered to pay crippling war damages, which became a source of deep national resentment that fuelled Nazi propaganda and is seen as a key underlying cause of World War II.

Germany was divided by the Danzig corridor which isolated eastern Prussia from the rest of the country. Berlin lost 15 per cent of its territory and 10 per cent of its population.

The regions of Alsace and Lorraine were returned to France and the Saarland region in Germany was placed under an international mandate for 15 years.

The treaty was never ratified by the United States Senate.

Saint-Germain-en-Laye: Sept 10, 1919

This treaty established peace between the Allies and defeated Austria, and consecrated the dismantlement of the Austro-Hungarian empire into several smaller states, a source of many future tensions.

Signed at the castle of Saint-Germain-en-Laye outside Paris, it recognised the creation of Czechoslovakia and the merger of southern Slav states that were to become Yugoslavia.

Romania expanded to include Transylvania and Bessarabia, and Poland was granted land occupied by Austria and Germany.

This left a separate Austria with 6.5 million inhabitants and Hungary with eight million.

Neuilly: Nov 27, 1919

Allied powers signed this treaty with Bulgaria, which had joined forces with Germany in 1915, in the Paris suburb of Neuilly.

It saw Sofia lose land to the newly formed Yugoslavia, to Romania and to Greece, leaving Bulgaria without direct access to the Aegean Sea.

Trianon: June 4, 1920

The Treaty of Trianon was concluded at Versailles between the Allies and Hungary, which lost two-thirds of the land it controlled as part of the Austro-Hungarian empire.

It meant that three million ethnic Hungarians found themselves living outside their country, for the most part in Romania.

Two with Turkey

Treaties signed with Turkey led to the dissolution of the once-mighty Ottoman empire and defined most of the borders of modern-day Turkey.

The first was inked at Sevres, in France, on Aug 10, 1920. Its terms led to Turkey's war of independence and a conflict with Greece, and was superseded by the Lausanne treaty on July 24, 1923, that ended the conflict.

Signed for the Allies by Britain, France and Italy, the treaties saw the country lose its Arab possessions, with Britain receiving a mandate for Palestine and Mesopotamia, and France awarded one for Lebanon and Syria.

Almost 1.3 million ethnic Greeks were forced to leave Asia Minor and about 500,000 Turks left Greece.