SINGAPORE - Singapore was one of five major biological warfare centres operated by the Japanese during World War II (WWII).
Laboratories here set up by Japan's biological warfare Unit 731 not only bred rat fleas and used them to spread the bubonic plague to enemies, but also cultivated smallpox virus, malaria parasite and tetanus bacteria.
Already a subscriber? Log in
Read the full story and more at $9.90/month
Get exclusive reports and insights with more than 500 subscriber-only articles every month
ST One Digital
$9.90/month
No contract
ST app access on 1 mobile device
Unlock these benefits
All subscriber-only content on ST app and straitstimes.com
Easy access any time via ST app on 1 mobile device
E-paper with 2-week archive so you won't miss out on content that matters to you