SAN FRANCISCO (BLOOMBERG) -Turmoil at Uber Technologies continues as longtime backer and director Bill Gurley is leaving the company's board following the resignation of chief executive officer Travis Kalanick.

Mr Gurley will hand the board seat to Mr Matt Cohler, his partner at Benchmark, a spokesman for the venture capital firm said. This will be the third change to Uber's board in recent weeks. TPG's David Bonderman was replaced as director by his colleague David Trujillo on Wednesday. Mr Bonderman resigned after he made a sexist remark to fellow board member Arianna Huffington at a company all-hands meeting meant to address Uber's flawed corporate culture.

Nestle executive Wan Ling Martello joined the board earlier this month.

Mr Gurley was one of Uber's earliest investors and biggest fans. He played a key role on the board over the years, helping to recruit many of the company's executives. But in private Mr Gurley became a vocal critic of Mr Kalanick in recent months, people familiar with the matter said.

The start-up has been in upheaval for most of this year, following rider protests over the company's ties to the Trump administration and an investigation into Uber's corporate culture conducted by former US attorney-general Eric Holder.

While Mr Gurley's criticism of Mr Kalanick endeared the venture capitalist to some, it also created enemies at Uber, people close to the company said. Mr Emil Michael, Mr Kalanick's chief confidant, accused Mr Gurley of putting his own reputation over the company's, two people close to Mr Michael told Bloomberg. Mr Michael was ousted last week.